Haplodrassus

Haplodrassus
	H. signifer
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Araneae
Infraorder:	Araneomorphae
Family:	Gnaphosidae
Genus:	Haplodrassus; Chamberlin, 1922
Type species
	H. hiemalis; (Emerton, 1909)
Species
	79, see text
Synonyms
	Tuvadrassus Marusik & Logunov, 1995;

Haplodrassus is a genus of ground spiders that was first described by R. V. Chamberlin in 1922.^[3] They range from 3 to 10 millimetres (0.12 to 0.39 in). H. signifer is the most widespread species, found across North America except for Alaska and northern Canada.^[4]

Species

As of May 2019^[update] it contains seventy-nine species:^[1]

H. aenus Thaler, 1984 – Switzerland, Austria
H. alexeevi Ponomarev & Shmatko, 2017 – Russia (Europe)
H. ambalaensis Gajbe, 1992 – India
H. atarot Levy, 2004 – Israel
H. belgeri Ovtsharenko & Marusik, 1988 – Russia (South to north-east Siberia, Far East)
H. bengalensis Gajbe, 1992 – India
H. bicornis (Emerton, 1909) – USA, Canada
H. bohemicus Miller & Buchar, 1977 – Czech Rep., Macedonia, Greece, Ukraine?, Russia (Europe, Caucasus)?
H. canariensis Schmidt, 1977 – Canary Is.
H. caspius Ponomarev & Belosludtsev, 2008 – Russia (Europe, Caucasus), Azerbaijan, Iran, Kazakhstan
H. caucasius Ponomarev & Dvadnenko, 2013 – Caucasus (Russia, Georgia)
H. chamberlini Platnick & Shadab, 1975 – North America
H. chotanagpurensis Gajbe, 1987 – India
H. cognatus (Westring, 1861) – Europe, Russia (Europe to Far East), Caucasus, Kazakhstan, Japan
- Haplodrassus c. ermolajewi Lohmander, 1942 – Russia (West Siberia)
H. concertor (Simon, 1878) – France
H. crassipes (Lucas, 1846) – Morocco, Algeria
H. creticus (Roewer, 1928) – Greece (Crete)
H. dalmatensis (L. Koch, 1866) – Europe, North Africa, Turkey, Middle East, Russia (Europe) to Central Asia
- Haplodrassus d. pictus (Thorell, 1875) – Spain, Madeira
H. dentatus Xu & Song, 1987 – China
H. dentifer Bosmans & Abrous, 2018 – Morocco, Algeria, Tunisia, Spain
H. deserticola Schmidt & Krause, 1996 – Canary Is.
H. dixiensis Chamberlin & Woodbury, 1929 – USA
H. dumdumensis Tikader, 1982 – India
H. eunis Chamberlin, 1922 – USA, Canada
H. hatsushibai Kamura, 2007 – Japan
H. hiemalis (Emerton, 1909) (type) – North America, Russia (Europe to Far East)
H. huarong Yin & Bao, 2012 – China
H. hunanensis Yin & Bao, 2012 – China
H. ibericus Melic, Silva & Barrientos, 2016 – Portugal, Spain
H. invalidus (O. Pickard-Cambridge, 1872) – Egypt, Cyprus, Turkey, Israel, Azerbaijan, Greece?
H. ivlievi Ponomarev, 2015 – Russia (Europe)
H. jacobi Gajbe, 1992 – India
H. kanenoi Kamura, 1995 – Japan
H. kulczynskii Lohmander, 1942 – Europe, Turkey, Russia (Europe to Far East), China, Korea
H. lilliputanus Levy, 2004 – Israel
H. longivulva Bosmans & Hervé, 2018 – Morocco, Algeria
H. lyndae Abrous & Bosmans, 2018 – Morocco, Algeria, Spain
H. macellinus (Thorell, 1871) – France, Italy, Portugal?, Spain?
H. maculatus (Banks, 1904) – USA, Mexico
H. mayumiae Kamura, 2007 – Korea, Japan
H. mediterraneus Levy, 2004 – Turkey, Syria, Lebanon, Israel, Jordan
H. mimus Chamberlin, 1922 – USA
H. minor (O. Pickard-Cambridge, 1879) – Europe, Turkey
H. moderatus (Kulczyński, 1897) – Europe, Russia (Europe to Far East), China
H. montanus Paik & Sohn, 1984 – Russia (Far East), China, Korea
H. morosus (O. Pickard-Cambridge, 1872) – Greece, Turkey, Israel
H. nigroscriptus (Simon, 1909) – Morocco
H. nojimai Kamura, 2007 – Japan
H. omissus (O. Pickard-Cambridge, 1872) – Canary Is., Morocco, Mediterranean
H. orientalis (L. Koch, 1866) – Bulgaria,^[5] Greece, Ukraine, Russia (Europe), Kazakhstan
H. ovatus Bosmans & Hervé, 2018 – Algeria, Tunisia
H. ovtchinnikovi Ponomarev, 2008 – Turkey, Iran, Kazakhstan
H. paramecus Zhang, Song & Zhu, 2001 – China
H. pargongsanensis Paik, 1992 – Korea
H. ponomarevi Kovblyuk & Seyyar, 2009 – Greece, Turkey
H. pseudosignifer Marusik, Hippa & Koponen, 1996 – Ukraine, Russia (Europe to Central Asia), Iran
H. pugnans (Simon, 1880) – Israel, Russia (Europe to Far East), China, Japan
H. reginae Schmidt & Krause, 1998 – Cape Verde Is.
H. rhodanicus (Simon, 1914) – Portugal, Spain, France, Italy (Sardinia), Tunisia
H. rufipes (Lucas, 1846) – Morocco, Algeria, Tunisia, Portugal, Spain, France, Italy
H. rufus (Savelyeva, 1972) – Kazakhstan
H. rugosus Tuneva, 2004 – Kazakhstan
H. sataraensis Tikader & Gajbe, 1977 – India
H. securifer Bosmans & Abrous, 2018 – Morocco, Algeria, Tunisia, Portugal, Spain, France, Italy, Belgium
H. signifer (C. L. Koch, 1839) – North America, Europe, North Africa, Turkey, Israel, Caucasus, Russia (Europe to Far East), Central Asia, China, Korea
H. silvestris (Blackwall, 1833) – Europe, Turkey, Caucasus
H. soerenseni (Strand, 1900) – Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, China
H. stuxbergi (L. Koch, 1879) – Russia (West to Middle Siberia)
H. taepaikensis Paik, 1992 – Russia (South Siberia, Far East), Korea
H. taibo (Chamberlin, 1919) – USA
H. tegulatus (Schenkel, 1963) – Russia (South Siberia), China
H. tehriensis Tikader & Gajbe, 1977 – India
H. triangularis Bosmans, 2018 – Morocco, Tunisia
H. typhon (Simon, 1878) – Algeria, Tunisia, Portugal, Spain, France, Italy (Sardinia)
H. umbratilis (L. Koch, 1866) – Europe to Kazakhstan
- Haplodrassus u. gothicus Lohmander, 1942 – Sweden
H. vastus (Hu, 1989) – China

References

^ ^a ^b ^c "Gen. Haplodrassus Chamberlin, 1922". World Spider Catalog Version 20.0. Natural History Museum Bern. 2019. doi:10.24436/2. Retrieved 2019-06-04.
^ Murphy, J. (2007). Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambridgeshire. p. 9.
^ Chamberlin, R. V. (1922). "The North American spiders of the family Gnaphosidae". Proceedings of the Biological Society of Washington. 35: 145–172.
^ "Genus Haplodrassus". BugGuide. Retrieved 2019-06-04.
^ Naumova, Maria; Blagoev, Gergin; Deltshev, Christo (June 2021). "Fifty spider species new to the Bulgarian fauna, with a review of some dubious species (Arachnida: Araneae)". Zootaxa. 4984 (1): 228–257. Retrieved 6 June 2024.

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[NMBE-1] "Gen. Haplodrassus Chamberlin, 1922". World Spider Catalog Version 20.0. Natural History Museum Bern. 2019. doi:10.24436/2. Retrieved 2019-06-04.

[Murp2007-2] Murphy, J. (2007). Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambridgeshire. p. 9.

[Cham1922-3] Chamberlin, R. V. (1922). "The North American spiders of the family Gnaphosidae". Proceedings of the Biological Society of Washington. 35: 145–172.

[bugguide-4] "Genus Haplodrassus". BugGuide. Retrieved 2019-06-04.

[5] Naumova, Maria; Blagoev, Gergin; Deltshev, Christo (June 2021). "Fifty spider species new to the Bulgarian fauna, with a review of some dubious species (Arachnida: Araneae)". Zootaxa. 4984 (1): 228–257. Retrieved 6 June 2024.

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