Isotelus

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Isotelus
Temporal range: Middle Ordovician–Late Ordovician
Isotelus gigas fossil at the Smithsonian National Museum of Natural History
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Asaphida
Family: Asaphidae
Genus: Isotelus
Dekay, 1824
Species

See text

Isotelus is a genus of asaphid trilobites from the Middle and Late Ordovician Period, fairly common in the northeastern United States, northwest Manitoba, southwestern Quebec and southeastern Ontario.

Gigantism[edit]

Cast of the I. rex holotype specimen, the largest complete trilobite specimen ever found.[1]

The mostly complete holotype specimen of Isotelus rex, from Churchill River Group, Churchill, Manitoba, is the largest complete trilobite ever found. Discovered by Dave Rudkin (Royal Ontario Museum), Robert Elias (University of Manitoba), Graham Young (Manitoba Museum) and Edward Dobrzanske (Manitoba Museum) in 1999, it measures 720 millimetres (28 in) in length, 400 millimetres (16 in) in maximum width (across the cephalon) and 70 millimetres (3 in) in maximum height (at the posterior midpoint of the cephalon).[2][3][4] It was probably exceeded in size only by Hungioides bohemicus, in which the specimens probably exceeding I. rex in size are only known from partial remains.[5]

Being found in Manitoba, which was equatorial during the Ordovician, I. rex is a dramatic example of low-latitude gigantism, contrasting with many modern marine benthic arthropods which adhere to Bergmann's rule of polar gigantism.[6]

Species[edit]

A number of species have been described:

  • I. aktchokensis Weber, 1948
  • I. bradleyi Amati, 2014
  • I. copenhagenensis Ross, Jr. & Shaw 1972
  • I. frognoensis Owen, 1981
  • I. gigas Dekay, 1824
  • I. harrisi Raymond, 1905 (synonym I. platymarginatus Raymond, 1910)
  • I. iowensis Owen, 1852
  • I. kimmswickensis Bradley, 1930
  • I. maximus Locke, 1838
  • I. megistos Locke, 1842
  • I. parvirugosus Chatterton & Ludvigsen, 1976
  • I. skapaneidos Amati, 2014
  • I. susae Whitfield, 1882
  • I. violaensis Amati, 2014
  • I. walcotti Walcott, 1918 (synonym I. planus De Mott, 1963)

Many specimens have been found in Ohio, where Isotelus is the State Invertebrate Fossil.[7][8]

References[edit]

  1. ^ Rudkin, David M.; Young, Graham A.; Elias, Robert J.; Dobrzanski, Edward P. (2003-01). "THE WORLD'S BIGGEST TRILOBITE—ISOTELUS REX NEW SPECIES FROM THE UPPER ORDOVICIAN OF NORTHERN MANITOBA, CANADA". Journal of Paleontology. 77 (1): 99–112. doi:10.1666/0022-3360(2003)077<0099:TWBTIR>2.0.CO;2. ISSN 0022-3360. {{cite journal}}: Check date values in: |date= (help)
  2. ^ Rudkin, D.A.; Young, G.A.; Elias, R.J.; Dobrzanske, E.P. (2003). "The World's biggest Trilobite: Isotelus rex new species from the Upper Ordovician of northern Manitoba, Canada". Journal of Paleontology. 70 (1): 99–112. doi:10.1666/0022-3360(2003)077<0099:TWBTIR>2.0.CO;2. ISSN 0022-3360.
  3. ^ S. M. Gon III (2005-08-17). "The World's Largest Trilobites".
  4. ^ P. Z. Myers (2005-04-25). "Isotelus rex, biggest trilobite ever". Archived from the original on 2007-11-12.
  5. ^ Gutiérrez-Marco, Juan C.; Sá, Artur A.; García-Bellido, Diego C.; Rábano, Isabel; Valério, Manuel (2009). "Giant trilobites and trilobite clusters from the Ordovician of Portugal". Geology. 37 (5): 443–446. Bibcode:2009Geo....37..443G. doi:10.1130/g25513a.1. ISSN 1943-2682.
  6. ^ Rudkin, David M.; Young, Graham A.; Elias, Robert J.; Dobrzanski, Edward P. (2003-01). "THE WORLD'S BIGGEST TRILOBITE—ISOTELUS REX NEW SPECIES FROM THE UPPER ORDOVICIAN OF NORTHERN MANITOBA, CANADA". Journal of Paleontology. 77 (1): 99–112. doi:10.1666/0022-3360(2003)077<0099:TWBTIR>2.0.CO;2. ISSN 0022-3360. {{cite journal}}: Check date values in: |date= (help)
  7. ^ "Ohio's State Fossil - Isotelus".
  8. ^ Babcock, L. E. (1996). "Chapter 8. Phylum Arthropoda, Class Trilobita". Fossils of Ohio. Columbus, Ohio: Ohio Division of Geological Survey, Bulletin 70. pp. 90–113. ISBN 0-931079-05-5.{{cite book}}: CS1 maint: date and year (link)