Phylinae

Phylinae
Atractotomus magnicornis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hemiptera
Suborder:	Heteroptera
Family:	Miridae
Subfamily:	Phylinae Douglas & Scott, 1865

Rhinocapsus vanduzeei

Phylinae is a subfamily of the plant bug family Miridae. Species of this family are found worldwide.

In research published in 2013, the subfamily Phylinae was reorganized. The tribe Auricillocorini is now considered a synonym of Hallodapini, and the tribe Pronotocrepini a synonym of the subtribe Cremnorrhina. New tribes were created under the new or resurrected names Decomiini, Exocarpocorini, Cremnorrhini, Nasocorini, and Semiini. The revised subfamily consists of nine tribes containing around 400 genera and over 2000 species.

Subfamily Phylinae Douglas & Scott, 1865

Tribe Cremnorrhinini Reuter, 1883 (sometimes spelled "Cremnorrhini")

Subtribe Coatonocapsina Schuh & Menard, 2013

Subtribe Cremnorrhina Reuter, 1883

Tribe Decomiini Schuh & Menard, 2013

Tribe Exaeretini Futon, 1875

Tribe Hallodapini Van Duzee, 1916

Tribe Leucophoropterini Schuh, 1974

Subtribe Leucophoropterina Schuh, 1974

Subtribe Tuxedoina Schuh & Menard, 2013

Tribe Nasocorini Reuter, 1883

Tribe Phylini Douglas & Scott, 1865

Subtribe Keltoniina Schuh & Menard, 2013

Subtribe Oncotylina Douglas & Scott, 1865

Subtribe Phylina Douglas & Scott, 1865

Tribe Pilophorini Douglas & Scott, 1876

Tribe Semiini Knight, 1923

Subtribe Exocarpocorina Schuh & Menard, 2013

Subtribe Semiina Knight, 1923

Genera

Phylinae contains the following genera:

Subfamily Phylinae Douglas & Scott, 1865

Tribe Cremnorrhinini Reuter, 1883

Subtribe Coatonocapsina Schuh & Menard, 2013

Genus Austropsallus Schuh, 1974

Genus Capecapsus Schuh, 1974

Genus Coatonocapsus Schuh, 1974

Genus Heterocapillus Wagner, 1960

Genus Parasciodema Poppius, 1914

Subtribe Cremnorrhinina Reuter, 1883

Genus Amblytylus Fieber, 1858

Genus Brachyceratocoris Knight, 1968

Genus Calidroides Schwartz, 2005

Genus Coquillettia Uhler, 1890

Genus Cremnorrhinus Reuter, 1880

Genus Dacota Uhler, 1872

Genus Denticulophallus Schuh, 1974

Genus Ethelastia Reuter, 1876

Genus Euderon Puton, 1888

Genus Excentricoris Carvalho, 1955

Genus Guentherocoris Schuh & Schwartz, 2004

Genus Harpocera Curtis, 1838

Genus Leutiola Wyniger, 2012

Genus Lopidodenus V. Putshkov, 1974

Genus Lopus Hahn, 1833

Genus Macrotylus Fieber, 1858

Genus Orectoderus Uhler, 1876

Genus Pachyxyphus Fieber, 1858

Genus Paralopus Wagner, 1957

Genus Pronotocrepis Knight, 1929

Genus Shendina Linnavuori, 1975

Genus Strophopoda Van Duzee, 1916

Genus Teleorhinus Uhler, 1890

Genus Ticua Wyniger, 2012

Genus Utopnia Reuter, 1881

Genus Zinjolopus Linnavuori, 1975

Tribe Decomiini Schuh & Menard, 2013

Genus Aurantiocoris Schuh & Schwartz, 2004

Genus Decomia Poppius, 1915

Genus Decomioides Schuh, 1984

Genus Malaysiamiris Schuh, 1984

Genus Malaysiamiroides Schuh, 1984

Genus Rubrocuneocoris Schuh, 1984

Tribe Exaeretini Futon, 1875

Genus Anonychiella Reuter, 1912

Genus Aphaenophyes Reuter, 1899

Genus Atractotomimus Kiritshenko, 1952

Genus Auchenocrepis Fieber, 1858

Genus Brendaphylus Yasunaga, 2013

Genus Camptotylus Fieber, 1860

Genus Camptozorus Kerzhner, 1996

Genus Chrysochnoodes Reuter, 1901

Genus Compsonannus Reuter, 1902

Genus Eumecotarsus Kerzhner, 1962

Genus Eurycranella Reuter, 1904

Genus Frotaphylus Carvalho, 1984

Genus Gonoporomiris Henry & Schuh, 2002

Genus Hadrophyes Puton, 1874

Genus Hyalopsallus Carvalho & Schaffner, 1973

Genus Megalodactylus Fieber, 1858

Genus Moissonia Reuter, 1894

Genus Opuna Kirkaldy, 1902

Genus Pastocoris Reuter, 1879

Genus Psallopsis Reuter, 1901

Genus Randallopsallus Yasunaga, 2013

Genus Tuponia Reuter, 1875

Genus Voruchiella Poppius, 1912

Genus Yotvata Linnavuori, 1964

Tribe Hallodapini Van Duzee, 1916

Genus Acrorrhinium Noualhier, 1895

Genus Aeolocoris Reuter, 1903

Genus Alloeomimus Reuter, 1910

Genus Artchawakomius Yasunaga, 2012

Genus Aspidacanthus Reuter, 1901

Genus Auricillocoris Schuh, 1984

Genus Azizus Distant, 1910

Genus Bibundiella Poppius, 1914

Genus Boopidella Reuter, 1907

Genus Carinogulus Schuh, 1974

Genus Chaetocapsus Poppius, 1914

Genus Clapmarius Distant, 1904

Genus Cleotomiris Schuh, 1984

Genus Cleotomiroides Schuh, 1984

Genus Cremnocephalus Eieber, 1860

Genus Cyrtopeltocoris Reuter, 1876

Genus Diocoris Kirkaldy, 1902

Genus Eminoculus Schuh, 1974

Genus Formicopsella Poppius, 1914

Genus Gampsodema Odhiambo, 1960

Genus Glaphyrocoris Reuter, 1903

Genus Hadrodapus Linnavuori, 1996

Genus Hallodapomimus Herczek, 2000

Genus Hallodapus Eieber, 1858

Genus Ifephylus Linnavuori, 1993

Genus Kapoetius Schmitz, 1969

Genus Laemocoris Reuter, 1879

Genus Leaina Linnavuori, 1974

Genus Leptomimus Herczek & Popov, 2010

Genus Lestonisca Carvalho, 1988

Genus Linacoris Carvalho, 1983

Genus Lissocapsus Bergroth, 1903

Genus Malgacheocoris Carvalho, 1952

Genus Mimocoris J. Scott, 1872

Genus Myombea China & Carvalho, 1951

Genus Myrmicomimus Reuter, 1881

Genus Neolaemocoris Wagner, 1975

Genus Omphalonotus Reuter, 1876

Genus Pangania Poppius, 1914

Genus Paralaemocoris Linnavuori, 1964

Genus Phoradendrepulus Polhemus & Polhemus, 1985

Genus Podullahas Schuh, 1984

Genus Pongocoris Linnavuori, 1975

Genus Ribautocapsus Wagner, 1962

Genus Ruwaba Linnavuori, 1975

Genus Skukuza Schuh, 1974

Genus Sohenus Distant, 1910

Genus Syngonus Bergroth, 1926

Genus Systellonotidea Poppius, 1914

Genus Systellonotopsis Poppius, 1914

Genus Systellonotus Pieber, 1858

Genus Trichophorella Reuter, 1905

Genus Trichophthalmocapsus Poppius, 1914

Genus Vitsikamiris Polhemus, 1994

Genus Wygomiris Schuh, 1984

Genus Zaratus Distant, 1909

Tribe Leucophoropterini Schuh, 1974

Subtribe Leucophoropterina Schuh, 1974

Genus Abuyogocoris Schuh, 1984

Genus Aitkenia Carvalho & Gross, 1982

Genus Arafuramiris Schuh, 1984

Genus Ausejanus Menard & Schuh, 2011

Genus Austrodapus Menard & Schuh, 2011

Genus Biromiris Schuh, 1984

Genus Blesingia Carvalho & Gross, 1982

Genus Collessicoris Carvalho & Gross, 1982

Genus Ctypomiris Schuh, 1984

Genus Gulacapsus Schuh, 1984

Genus Johnstonsonius Menard & Schuh, 2011

Genus Leucophoroptera Poppius, 1921

Genus Missanos Menard & Schuh, 2011

Genus Neaitkenia Menard & Schuh, 2011

Genus Neoleucophoroptera Menard & Schuh, 2011

Genus Papuamimus Schuh, 1984

Genus Papuamiroides Menard & Schuh, 2011

Genus Pseudohallodapocoris Schuh, 1984

Genus Sejanus Distant, 1910

Genus Solomonomimus Schuh, 1984

Genus Transleucophoroptera Menard & Schuh, 2011

Genus Trichocephalocapsus Schuh, 1984

Genus Waterhouseana Carvalho, 1973

Subtribe Tuxedoina Schuh & Menard, 2013

Genus Ephippiocoris Poppius, 1912

Genus Pseudophylus Yasunaga, 1999

Genus Tuxedo Schuh, 2001

Tribe Nasocorini Reuter, 1883

Genus Adenostomocoris Schuh & Schwartz, 2004

Genus Agrametra Buchanan-White, 1878

Genus Arctostaphylocoris Schuh & Schwartz, 2004

Genus Atomophora Reuter, 1879

Genus Atomoscelis Reuter, 1875

Genus Atractotomoidea Yasunaga, 1999

Genus Atractotomus Fieber, 1858

Genus Badezorus Distant, 1910

Genus Beckocoris Knight, 1968

Genus Bergmiris Carvalho, 1984

Genus Boopidocoris Reuter, 1879

Genus Caiganga Carvalho & Becker, 1957

Genus Camptotylidea Wagner, 1957

Genus Campylomma Reuter, 1878

Genus Chinacapsus Wagner, 1961

Genus Chlamydatus Curtis, 1833

Genus Chlamyopsallus Schwartz, 2005

Genus Hambletoniola Carvalho, 1954

Genus Helenocoris Schmitz, 1976

Genus Hirtopsallus Schmitz, 1976

Genus Insulopus Schmitz, 1976

Genus Karocris V. Putshkov, 1975

Genus Kasumiphylus Schwartz & Stonedahl, 2004

Genus Knightomiroides Stonedahl & Schwartz, 1996

Genus Lamprosthenarus Poppius, 1914

Genus Larinocerus

Genus Lattinophylus Schuh, 2008

Genus Lindbergopsallus Wagner, 1962

Genus Lopsallus Schmitz, 1976

Genus Maurodactylus Reuter, 1878

Genus Megalopsallus Knight, 1927

Genus Monosynamma J. Scott, 1864

Genus Naresthus Schmitz, 1976

Genus Nasocoris Reuter, 1879

Genus Neisopsallus Schmitz, 1976

Genus Neophylus Carvalho & Costa, 1992

Genus Nevadocoris Knight, 1968

Genus Nigrimiris Carvalho & Schaffner, 1973

Genus Nigrocapillocoris Wagner, 1973

Genus Oligobiella Reuter, 1885

Genus Orthopidea Reuter, 1899

Genus Phaxia Kerzhner, 1984

Genus Phoenicocoris Reuter, 1875

Genus Pinomiris Stonedahl & Schwartz, 1996

Genus Pruneocoris Schuh & Schwartz, 2004

Genus Psallomimus Wagner, 1951

Genus Rhinacloa Reuter, 1876

Genus Salicarus Kerzhner, 1962

Genus Solenoxyphus Reuter, 1875

Genus Spanagonicus Berg, 1883

Genus Squamophylus Carvalho & Costa, 1992

Genus Sthenaropsis Poppius, 1912

Genus Taeniophorus Linnavuori, 1952

Genus Tannerocoris Knight, 1970

Genus Tapuruyunus Carvalho, 1946

Genus Thymopsallus Linnavuori, 1975

Genus Tijucaphylus Carvalho & Costa, 1992

Genus Tunisiella Carapezza, 1997

Genus Voruchia Reuter, 1879

Tribe Phylini Douglas & Scott, 1865

Subtribe Keltoniina Schuh & Menard, 2013

Genus Keltonia Knight, 1966

Genus Pseudatomoscelis Poppius, 1911

Genus Reuteroscopus Kirkaldy, 1905

Genus Waupsallus Linnavuori, 1975

Subtribe Oncotylina Douglas & Scott, 1865

Genus Acrotelus Reuter, 1885

Genus Alloeotarsus Reuter, 1885

Genus Americodema T. Henry, 1999

Genus Angelopsallus Schuh, 2006

Genus Antepia Seidenstiicker, 1962

Genus Arizonapsallus Schuh, 2006

Genus Asciodema Reuter, 1878

Genus Bisulcopsallus Schuh, 2006

Genus Brachyarthrum Fieber, 1858

Genus Cariniocoris T. Henry, 1989

Genus Ceratopsallus Schuh, 2006

Genus Cercocarpopsallus Schuh, 2006

Genus Chlorillus Kerzhner, 1962

Genus Compsidolon Reuter, 1899

Genus Crassomiris Weirauch, 2006

Genus Damioscea Reuter, 1883

Genus Dasycapsus Poppius, 1912

Genus Europiella Reuter, 1909

Genus Eurycolpus Reuter, 1875

Genus Galbinocoris Weirauch, 2006

Genus Glaucopterum Wagner, 1963

Genus Hamatophylus Weirauch, 2006

Genus Hoplomachus Fieber, 1858

Genus Ihermocoris Puton, 1875

Genus Insulaphylus Weirauch, 2006

Genus Josifovius Konstantinov, 2008

Genus Kmentophylus Duwal, Yasunaga, & Lee, 2010

Genus Knightophylinia Schaffner, 1978

Genus Knightopiella Schuh, 2004

Genus Knightopsallus Schuh, 2006

Genus Leucodellus Reuter, 1906

Genus Lineatopsallus T. Henry, 1991

Genus Litoxenus Reuter, 1885

Genus Maculamiris Weirauch, 2006

Genus Malacotes Reuter, 1878

Genus Marrubiocoris Wagner, 1970

Genus Megalocoleus Reuter, 1890

Genus Moiseevichia Schuh, 2006

Genus Nanopsallus Wagner, 1952

Genus Neopsallus Schuh & Schwartz, 2004

Genus Occidentodema T. Henry, 1999

Genus Oligotylus Van Duzee, 1916

Genus Omocoris Lindberg, 1930

Genus Oncotylidea Wagner, 1965

Genus Oncotylus Fieber, 1858

Genus Opisthotaenia Reuter, 1901

Genus Oreocapsus Linnavuori, 1975

Genus Parachlorillus Wagner, 1963

Genus Parapsallus Wagner, 1952

Genus Paredrocoris Reuter, 1878

Genus Phaeochiton Kerzhner, 1964

Genus Phallospinophylus Weirauch, 2006

Genus Phyllopidea Knight, 1919

Genus Phymatopsallus Knight

Genus Piceophylus Schwartz & Schuh, 1999

Genus Pinophylus Schwartz & Schuh, 1999

Genus Placochilus Fieber, 1858

Genus Plagiognathus Fieber, 1858

Genus Plesiodema Reuter, 1875

Genus Pleuroxonotus Reuter, 1903

Genus Pronototropis Reuter, 1879

Genus Psallodema V. Putshkov, 1970

Genus Psallomorpha Duwal Yasunaga, & Lee, 2010

Genus Psallovius T. Henry

Genus Pygovepres Weirauch, 2006

Genus Quercophylus Weirauch, 2006

Genus Quernocoris Weirauch, 2006

Genus Ranzovius Distant, 1893

Genus Rhinocapsus Uhler, 1890

Genus Roburocoris Weirauch, 2009

Genus Rubellomiris Weirauch, 2006

Genus Rubeospineus Weirauch, 2006

Genus Sacculifer Kerzhner, 1959

Genus Salicopsallus Schuh, 2006

Genus Schaffneropsallus Schuh, 2006

Genus Stenoparia Fieber, 1870

Genus Sthenaropsidea Henry & Schuh, 2002

Genus Stictopsallus Schuh, 2006

Genus Stirophylus Eckerlein & Wagner, 1965

Genus Stoebea Schuh, 1974

Genus Tapirula Carapezza, 1997

Genus Tinicephalus Fieber, 1858

Genus Tragiscocoris Fieber, 1861

Genus Vanduzeephylus Schuh & Schwartz, 2004

Genus Vesperocoris Weirauch, 2006

Genus Viscacoris Weirauch, 2009

Genus Zophocnemis Kerzhner, 1962

Subtribe Phylina Douglas & Scott, 1865

Genus Adelphophylus Wagner, 1959

Genus Agraptocoris Reuter, 1903

Genus Alnopsallus Duwal, Yasunaga, & Lee, 2010

Genus Alvarengamiris Carvalho, 1991

Genus Amazonophilus Carvalho & Costa, 1993

Genus Anapsallus Odhiambo, 1960

Genus Anomalocornis Carvalho & Wygodzinsky, 1945

Genus Arlemiris Carvalho, 1984

Genus Bicurvicoris Carvalho & Schaffner, 1973

Genus Botocudomiris Carvalho, 1979

Genus Brachycranella Reuter, 1905

Genus Conostethus Fieber, 1858

Genus Crassicornus Carvalho, 1945

Genus Darectagela Linnavuori, 1975

Genus Darfuromma Linnavuori, 1975

Genus Dignaia Linnavuori, 1975

Genus Dominiquella Linnavuori, 1983

Genus Ectagela Schmidt, K., 1939

Genus Ellacapsus Yasunaga, 2013

Genus Eremophylus Yasunaga, 2001

Genus Farsiana Linnavuori, 1998

Genus Gediocoris Wagner, 1964

Genus Ghazalocoris Linnavuori, 1975

Genus Gressittocapsus Schuh, 1984

Genus Icodema Reuter, 1875

Genus Indatractus Linnavuori, 1975

Genus Izyaius Schwartz, 2006

Genus Juniperia Linnavuori, 1965

Genus Knightensis Schaffner, 1978

Genus Lalyocoris Linnavuori, 1993

Genus Lasiolabopella Schuh, 1974

Genus Lepidargyrus Muminov, 1962

Genus Lepidocapsus Poppius, 1914

Genus Leptoxanthus Reuter, 1905

Genus Liviopsallus Carapezza, 1982

Genus Mendozaphylus Carvalho & Carpintero, 1991

Genus Millerimiris Carvalho, 1951

Genus Mixtecamiris Carvalho & Schaffner, 1973

Genus Natalophylus Schuh, 1974

Genus Nubaia Linnavuori, 1975

Genus Orthonotus Stephens, 1829

Genus Parafulvius Carvalho, 1954

Genus Paravoruchia Wagner, 1959

Genus Phylus Hahn, 1831

Genus Plagiognathidea Poppius, 1914

Genus Platyscytisca Costa & Henry, 1999

Genus Platyscytus Reuter, 1907

Genus Porophoroptera Carvalho & Gross, 1982

Genus Psallus Fieber, 1858

Genus Roudairea Puton & Reuter, 1886

Genus Sasajiophylus Yasunaga, 2001

Genus Somalocoris Linnavuori, 1975

Genus Sthenarus Fieber, 1858

Genus Stibaromma Odhiambo, 1961

Genus Tibiopilus Carvalho & Costa, 1993

Genus Trevessa China, 1924

Genus Villaverdea Carvalho, 1990

Genus Widdringtoniola Schuh, 1974

Genus Zakanocoris V. Putshkov, 1970

Genus Zanchiophylus Duwal, Yasunaga, & Lee, 2010

Tribe Pilophorini Douglas & Scott, 1876

Genus Alepidiella Poppius, 1914

Genus Aloea Linnavuori, 1975

Genus Dilatops Weirauch, 2006

Genus Druthmarus Distant, 1909

Genus Ethatractus Linnavuori, 1975

Genus Hypseloecus Reuter, 1891

Genus Lasiolabops Poppius, 1914

Genus Neoambonea Schuh, 1974

Genus Parambonea Schuh, 1974

Genus Parasthenaridea Miller, 1937

Genus Pherolepis Kulik, 1968

Genus Pilophorus Hahn, 1826

Genus Pseudambonea Schuh, 1974

Genus Randallophorus Henry, 2013

Genus Spinolosus Zou, 1985

Genus Sthenaridea Reuter, 1885

Tribe Semiini Knight, 1923

Subtribe Exocarpocorina Schuh & Menard, 2013

Genus Ampimpacoris Weirauch & Schuh, 2011

Genus Ancoraphylus Weirauch, 2007

Genus Araucanophylus Carvalho, 1984

Genus Basileobius Eyles & Schuh, 2003

Genus Chiloephylus Carvalho, 1984

Genus Cyrtodiridius Eyles & Schuh, 2003

Genus Exocarpocoris Weirauch, 2007

Genus Gonzalezinus Carvalho, 1981

Genus Halormus Eyles & Schuh, 2003

Genus Harpagophylus Schuh & Weirauch, 2010

Genus Jiwarli Soto & Weirauch, 2007

Genus Leptidolon Reuter, 1904

Genus Mecenopa Eyles & Schuh, 2003

Genus Melaleucoides Schuh & Weirauch, 2010

Genus Pimeleocoris Eyles & Schuh, 2003

Genus Polyozus Eyles & Schuh, 2003

Genus Thryptomenomiris Schuh & Weirauch, 2010

Genus Wallabicoris Schuh & Pedraza, 2010

Genus Xiphoidellus Weirauch & Schuh, 2011

Genus Xiphoides Eyles & Schuh, 2003

Subtribe Semiina Knight, 1923

Genus Criocoris Fieber, 1858

Genus Hoplomachidea Reuter, 1909

Genus Karoocapsus Schuh, 1974

Genus Monocris V. Putshkov, 1974

Genus Monospatha Eyles & Schuh 2003

Genus Myrmicopsella Poppius, 1914

Genus Nicholia Knight, 1929

Genus Parapseudosthenarus Schuh, 1974

Genus Pseudosthenarus Poppius, 1914

Genus Rakula Odhiambo, 1967

Genus Semium Reuter, 1876

Genus Thoth Linnavuori, 1993

Genus Tytthus Fieber, 1864

References

• Duwal, R.; Yasunaga, T. & Lee, S. (2010). "Revision of the plant bug tribe Phylini from Nepal (Heteroptera: Miridae: Phylinae)". Entomologica Americana. 116 (1): 1–48. doi:10.1664/08-RA-010R.1. S2CID 86821766.
• Yasunaga, T. (2010). "Plant bugs of the tribe Phylini in Thailand (Heteroptera: Miridae: Phylinae), with descriptions of six new species from additional areas in tropical and subtropical Asia". Entomologica Americana. 116 (3): 50–92. doi:10.1664/10-RA-006.1. S2CID 84983192.