Portal:Amphibians
The Amphibian Portal
Amphibians are ectothermic, anamniotic, four-limbed vertebrate animals that constitute the class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all tetrapods excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds and mammals). All extant (living) amphibians belong to the monophyletic subclass Lissamphibia, with three living orders: Anura (frogs and toads), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their life cycle typically starts out as aquatic larvae with gills known as tadpoles, but some species have developed behavioural adaptations to bypass this.
Young amphibians generally undergo metamorphosis from an aquatic larval form with gills to an air-breathing adult form with lungs. Amphibians use their skin as a secondary respiratory interface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards, but unlike reptiles and other amniotes, require access to water bodies to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators to habitat conditions; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.
The earliest amphibians evolved in the Devonian period from tetrapodomorph sarcopterygians (lobe-finned fish with articulated limb-like fins) that evolved primitive lungs, which were helpful in adapting to dry land. They diversified and became ecologically dominant during the Carboniferous and Permian periods, but were later displaced in terrestrial environments by early reptiles and basal synapsids (predecessors of mammals). The origin of modern lissamphibians, which first appeared during the Early Triassic, around 250 million years ago, has long been contentious. The most popular hypothesis is that they likely originated from temnospondyls, the most diverse group of prehistoric amphibians, during the Permian period. Another hypothesis is that they emerged from lepospondyls. A fourth group of lissamphibians, the Albanerpetontidae, became extinct around 2 million years ago. (Full article...)
Selected frog article
The Alsodidae are a small family of frogs from South America between Patagonia and southern Brazil. It contains 30 species in three genera. This family, along with several other families, used to be included in the family Leptodactylidae. It was then a subfamily in the family Cycloramphidae, before being recognized as a family first in 2011. (Full article...)
Selected salamander article
The Caudata are a group of amphibians containing the extant salamanders (the order Urodela) and all extinct species of amphibians more closely related to salamanders than to frogs. They are typically characterized by a superficially lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults.
Disagreement exists between different authorities as to the definition of the terms "Caudata" and "Urodela". Some maintain that Urodela should be restricted to the crown group, with Caudata being used for the total group. Others restrict the name Caudata to the crown group and use Urodela for the total group. The former approach seems to be most widely adopted and is used in this article. (Full article...)
List of selected salamander articles
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Did you know? –
- ...that the Cheat Mountain salamander of West Virginia is the only vertebrate known to be unique to that state?
- ... that the Permian amphibian Trimerorhachis either brooded young in its mouth or ate them?
- ... that the Cumberland Dusky Salamander is believed to hybridize with the Ocoee Salamander?
- ... that female European cave salamanders do not normally eat while they guard their eggs for six months or more?
- ... that in hot weather, the eastern spadefoot toad (pictured) may aestivate in the deep burrow it has dug?
Selected amphibian type
A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (coming from the Ancient Greek ἀνούρα, literally 'without tail'). The oldest fossil "proto-frog" Triadobatrachus is known from the Early Triassic of Madagascar (250 million years ago), but molecular clock dating suggests their split from other amphibians may extend further back to the Permian, 265 million years ago. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest. Frogs account for around 88% of extant amphibian species. They are also one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from taxonomy or evolutionary history.
An adult frog has a stout body, protruding eyes, anteriorly-attached tongue, limbs folded underneath, and no tail (the tail of tailed frogs is an extension of the male cloaca). Frogs have glandular skin, with secretions ranging from distasteful to toxic. Their skin varies in colour from well-camouflaged dappled brown, grey and green to vivid patterns of bright red or yellow and black to show toxicity and ward off predators. Adult frogs live in fresh water and on dry land; some species are adapted for living underground or in trees.
Frogs typically lay their eggs in the water. The eggs hatch into aquatic larvae called tadpoles that have tails and internal gills. They have highly specialised rasping mouth parts suitable for herbivorous, omnivorous or planktivorous diets. The life cycle is completed when they metamorphose into adults. A few species deposit eggs on land or bypass the tadpole stage. Adult frogs generally have a carnivorous diet consisting of small invertebrates, but omnivorous species exist and a few feed on plant matter. Frog skin has a rich microbiome which is important to their health. Frogs are extremely efficient at converting what they eat into body mass. They are an important food source for predators and part of the food web dynamics of many of the world's ecosystems. The skin is semi-permeable, making them susceptible to dehydration, so they either live in moist places or have special adaptations to deal with dry habitats. Frogs produce a wide range of vocalisations, particularly in their breeding season, and exhibit many different kinds of complex behaviors to attract mates, to fend off predators and to generally survive. (Full article...)
List of selected amphibian type articles
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Selected images
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The Eastern American Toad (Bufo americanus americanus) is a common subspecies of toad found throughout the eastern United States and Canada. It typically grows to 5–9 cm (2.0–3.5 in), with varying skin color and pattern depending on its environment. Its skin secretes bufotoxin, which is mildly poisonous to humans. -
Photo: Benny TrappAn Italian tree frog (Hyla intermedia) with an inflated vocal sac. Vocal sacs, flexible membranes of skin possessed by most male frogs, are generally used to amplify mating or advertisement calls. However, some species in the family Rhinodermatidae may also use the sac to store or transport their tadpoles. -
Photograph: Richard Bartz -
Photograph: JJ HarrisonThe southern brown tree frog (Litoria ewingii) is a species of tree frog native to Australia found in a wide range of habitats. Reaching 45 millimetres (1.8 in), this species is generally brown, but green and green-striped morphs are also recorded. -
Photo credit: Patrick CoinA terrestrial subadult Eastern newt or red eft (Notophthalmus viridescens). Salamanders of the family Salamandridae with aquatic adult stages are called newts. Some newts, including the Eastern newt, have a juvenile terrestrial stage called the eft. The red eft has bright aposematic coloration to warn predators of its highly toxic skin. -
Photograph: H. KrispThe phantasmal poison frog (Epipedobates tricolor) is a species of poison dart frog endemic to Ecuador. First described by George Albert Boulenger in 1899, this endangered species is found only in rainforests on the slopes of the Andes. -
Photograph: Benny TrappThe Iberian midwife toad (Alytes cisternasii) is a species of frog in the family Alytidae found in Portugal and western Spain. Its natural habitats are temperate forests and shrubland, Mediterranean-type shrubby vegetation, intermittent rivers and freshwater marshes, pastureland, and aquaculture ponds. The species is affected by habitat loss, and the IUCN lists it as being "Near Threatened". -
Photograph: Benny TrappThe Spanish painted frog (Discoglossus jeanneae) is a species of frog in the family Alytidae. Endemic to Spain, it mostly lives in open areas, pine groves and shrublands. It feeds mostly on insects and worms. -
Photograph credit: RushenbWallace's flying frog (Rhacophorus nigropalmatus) is a moss frog found in tropical southeastern Asia. It is named after the British naturalist Alfred R. Wallace, who collected the first known specimen of the species. It lives almost exclusively in trees, and when threatened, or in search of prey, will leap from a branch and splay its four webbed feet; the membranes between its toes and the loose skin flaps on its sides catch the air as it falls, helping it to glide. This individual was photographed in Khao Sok National Park, Thailand. -
Photo: Marie-Lan NguyenThe marsh frog (Pelophylax ridibundus) is a species of true frog and the largest frog native to Europe; females of this sexually dimorphic species may be up to 17 centimetres (6.7 in) long. The marsh frog feeds mainly on insects, but it also eats smaller amphibians, fish, and rodents. -
Photo credit: Fir0002The Eastern Banjo Frog (Limnodynastes dumerilli) is native to eastern Australia and has been introduced to New Zealand. The frog is also commonly called the pobblebonk after its distinctive "bonk" call, which is likened to a banjo string being plucked. Adults are roughly seven to eight cm long. -
The white-lipped tree frog (Litoria infrafrenata) is the world's largest tree frog. This species is native to the rainforests of Northern Queensland, New Guinea, the Bismarck Islands and the Admiralty Islands. It can reach a length of over 13 cm (5 in), with females growing larger than males. The lower lip has a distinctive white stripe, giving this species its common name. -
Photo credit: Derek RamseyThe American Green Frog (Rana clamitans) is native to the eastern half of the United States and Canada. Green frogs live wherever there is shallow freshwater: ponds, road-side ditches, lakes, swamps, streams and brooks. The green frog is abundant wherever it occurs. -
The fire salamander (Salamandra salamandra) is a European salamander species with a long lifespan. These nocturnal animals generally eat various insects, spiders, earthworms and slugs, but they also occasionally eat newts and young frogs. -
Photo: QuartlDendrobates azureus, commonly known as the blue poison dart frog or okopipi, is a poison dart frog found in the forests surrounded by the Sipaliwini Savanna in southern Suriname. Authorities have recently treated it as a variant of Dendrobates tinctorius rather than a distinct species as before.
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Selected toad article
The cane toad (Rhinella marina), also known as the giant neotropical toad or marine toad, is a large, terrestrial true toad native to South and mainland Central America, but which has been introduced to various islands throughout Oceania and the Caribbean, as well as Northern Australia. It is a member of the genus Rhinella, which includes many true toad species found throughout Central and South America, but it was formerly assigned to the genus Bufo.
A fossil toad (specimen UCMP 41159) from the La Venta fauna of the late Miocene in Colombia is morphologically indistinguishable from modern cane toads from northern South America. It was discovered in a floodplain deposit, which suggests the R. marina habitat preferences have long been for open areas. The cane toad is a prolific breeder; females lay single-clump spawns with thousands of eggs. Its reproductive success is partly because of opportunistic feeding: it has a diet, unusual among anurans, of both dead and living matter. Adults average 10–15 cm (4–6 in) in length; the largest recorded specimen had a snout-vent length of 24 cm (9.4 in).
The cane toad has poison glands, and the tadpoles are highly toxic to most animals if ingested. Its toxic skin can kill many animals, both wild and domesticated, and cane toads are particularly dangerous to dogs. Because of its voracious appetite, the cane toad has been introduced to many regions of the Pacific and the Caribbean islands as a method of agricultural pest control. The common name of the species is derived from its use against the cane beetle (Dermolepida albohirtum), which damages sugar cane. The cane toad is now considered a pest and an invasive species in many of its introduced regions. The 1988 film Cane Toads: An Unnatural History documented the trials and tribulations of the introduction of cane toads in Australia. (Full article...)
Selected caecilian article
Chthonerpeton is a genus of semiaquatic amphibians in the family Typhlonectidae. They occur in South America east of the Andes. (Full article...)
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