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The Silurian Portal

Introduction

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Selected article on the Silurian world and its legacies

A tunicate is a marine invertebrate animal, a member of the subphylum Tunicata which is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords. Some tunicates live as solitary individuals but others replicate by budding and become colonies, each unit being known as a zooid. They are marine filter feeders with a water-filled, sac-like body structure and two tubular openings, known as siphons, through which they draw in and expel water. During their respiration and feeding they take in water through the incurrent (or inhalant) siphon and expel the filtered water through the excurrent (or exhalant) siphon. Most adult tunicates are sessile and are permanently attached to rocks or other hard surfaces on the ocean floor; others such as salps, doliolids and pyrosomes swim in the pelagic zone of the sea as adults. Various species are commonly known as sea squirts, sea pork, sea liver or sea tulips.

The Tunicata first appear in the fossil record in the early Cambrian period. Despite their simple appearance and very different adult form, their close relationship to the vertebrates is shown by the fact that during their mobile larval stage, they possess a notochord or stiffening rod and resemble a tadpole. Their name derives from their unique outer covering or "tunic" which is formed from proteins and carbohydrates and acts as an exoskeleton. In some species it is thin, translucent and gelatinous while in others it is thick, tough and stiff. (see more...)

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Selected article on the Silurian in human science, culture and economics

The history of paleontology traces the history of the effort to study the fossil record left behind by ancient life forms. Although fossils had been studied by scholars since ancient times, the nature of fossils and their relationship to life in the past became better understood during the 17th and 18th centuries. At the end of the 18th century the work of Georges Cuvier ended a long running debate about the reality of extinction and led to the emergence of paleontology as a scientific discipline.

The first half of the 19th century saw paleontological activity become increasingly well organized. This contributed to a rapid increase in knowledge about the history of life on Earth, and progress towards definition of the geologic time scale. As knowledge of life's history continued to improve, it became increasingly obvious that there had been some kind of successive order to the development of life. After Charles Darwin published Origin of Species in 1859, much of the focus of paleontology shifted to understanding evolutionary paths.

The last half of the 19th century saw a tremendous expansion in paleontological activity, especially in North America. The trend continued in the 20th century with additional regions of the Earth being opened to systematic fossil collection, as demonstrated by a series of important discoveries in China near the end of the 20th century. There was also a renewed interest in the Cambrian explosion that saw the development of the body plans of most animal phyla. (see more...)

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Fossils of the Silurian age from the 4th edition of Meyers Konversationslexikon (1885–90).
Photo credit: Bibliographisches Institut

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Did you know?

... that a fossil of the extinct sea scorpion Eurypterus (restoration pictured) was once thought to be a catfish?
... that the Paleozoic monoplacophoran Pilina unguis was twice as large as any known living monoplacophoran?
... that the extinct trilobite Bumastus (artist's rendition pictured) was named after its resemblance to large grapes?
...that Ornatifilum is likely to be the oldest known fossil fungus?
...that the Silurian Lau extinction event caused the Earth to temporarily return to a condition similar to how it was before multicellular life evolved?